Karl Von Baer
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Karl Ernst Ritter von Baer Edler von Huthorn ( – ) was a
Baltic German Baltic Germans (german: Deutsch-Balten or , later ) were ethnic German inhabitants of the eastern shores of the Baltic Sea, in what today are Estonia and Latvia. Since their coerced resettlement in 1939, Baltic Germans have markedly declined ...
scientist and explorer. Baer was a naturalist,
biologist A biologist is a scientist who conducts research in biology. Biologists are interested in studying life on Earth, whether it is an individual Cell (biology), cell, a multicellular organism, or a Community (ecology), community of Biological inter ...
,
geologist A geologist is a scientist who studies the solid, liquid, and gaseous matter that constitutes Earth and other terrestrial planets, as well as the processes that shape them. Geologists usually study geology, earth science, or geophysics, althou ...
,
meteorologist A meteorologist is a scientist who studies and works in the field of meteorology aiming to understand or predict Earth's atmospheric phenomena including the weather. Those who study meteorological phenomena are meteorologists in research, while t ...
,
geographer A geographer is a physical scientist, social scientist or humanist whose area of study is geography, the study of Earth's natural environment and human society, including how society and nature interacts. The Greek prefix "geo" means "earth" a ...
, and is considered a, or the, founding father of
embryology Embryology (from Greek ἔμβρυον, ''embryon'', "the unborn, embryo"; and -λογία, '' -logia'') is the branch of animal biology that studies the prenatal development of gametes (sex cells), fertilization, and development of embryos and ...
. He was a member of the
Russian Russian(s) refers to anything related to Russia, including: *Russians (, ''russkiye''), an ethnic group of the East Slavic peoples, primarily living in Russia and neighboring countries *Rossiyane (), Russian language term for all citizens and peo ...
Academy of Sciences An academy of sciences is a type of learned society or academy (as special scientific institution) dedicated to sciences that may or may not be state funded. Some state funded academies are tuned into national or royal (in case of the Unite ...
, a co-founder of the
Russian Geographical Society The Russian Geographical Society (russian: Ру́сское географи́ческое о́бщество «РГО»), or RGO, is a learned society based in Saint Petersburg, Russia. It promotes geography, exploration and nature protection wi ...
, and the first president of the
Russian Russian(s) refers to anything related to Russia, including: *Russians (, ''russkiye''), an ethnic group of the East Slavic peoples, primarily living in Russia and neighboring countries *Rossiyane (), Russian language term for all citizens and peo ...
Entomological Entomology () is the scientific study of insects, a branch of zoology. In the past the term "insect" was less specific, and historically the definition of entomology would also include the study of animals in other arthropod groups, such as arach ...
Society A society is a group of individuals involved in persistent social interaction, or a large social group sharing the same spatial or social territory, typically subject to the same political authority and dominant cultural expectations. Socie ...
, making him one of the most distinguished Baltic German scientists.


Life

Karl Ernst von Baer was born into the Baltic German noble Baer family ( et) in the Piep Manor ( et), Jerwen County,
Governorate of Estonia The Governorate of Estonia, also known as the Governorate of Esthonia (Pre-reformed rus, Эстля́ндская губе́рнія, r=Estlyandskaya guberniya); et, Eestimaa kubermang was a governorate in the Baltic region, along with th ...
(in present-day
Lääne-Viru County Lääne-Viru County ( et, Lääne-Viru maakond or ''Lääne-Virumaa'') is one of 15 counties of Estonia. It is in northern Estonia, on the south coast of the Gulf of Finland. In Estonian, ''lääne'' means western and ''ida'' means east or eastern ...
,
Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, a ...
), as a
knight A knight is a person granted an honorary title of knighthood by a head of state (including the Pope) or representative for service to the monarch, the church or the country, especially in a military capacity. Knighthood finds origins in the Gr ...
by birthright. His patrilineal ancestors were of
Westphalia Westphalia (; german: Westfalen ; nds, Westfalen ) is a region of northwestern Germany and one of the three historic parts of the state of North Rhine-Westphalia. It has an area of and 7.9 million inhabitants. The territory of the regio ...
n origin and originated in
Osnabrück Osnabrück (; wep, Ossenbrügge; archaic ''Osnaburg'') is a city in the German state of Lower Saxony. It is situated on the river Hase in a valley penned between the Wiehen Hills and the northern tip of the Teutoburg Forest. With a population ...
. He spent his early childhood at
Lasila manor Lasila is a village in Rakvere Parish, Lääne-Viru County, in northeastern Estonia. Lasila manor Lasila estate dates from the end of the 17th century. The current building was erected in 1862 in a romantic, Gothic Revival architecture, neo-Gothi ...
, Estonia. He was educated at the Knight and Cathedral School in
Reval Tallinn () is the most populous and capital city of Estonia. Situated on a bay in north Estonia, on the shore of the Gulf of Finland of the Baltic Sea, Tallinn has a population of 437,811 (as of 2022) and administratively lies in the Harju ''m ...
(Tallinn) and the
Imperial University of Dorpat The University of Tartu (UT; et, Tartu Ülikool; la, Universitas Tartuensis) is a university in the city of Tartu in Estonia. It is the national university of Estonia. It is the only classical university in the country, and also its biggest ...
(
Tartu Tartu is the second largest city in Estonia after the Northern European country's political and financial capital, Tallinn. Tartu has a population of 91,407 (as of 2021). It is southeast of Tallinn and 245 kilometres (152 miles) northeast of ...
). In 1812, during his tenure at the university, he was sent to
Riga Riga (; lv, Rīga , liv, Rīgõ) is the capital and largest city of Latvia and is home to 605,802 inhabitants which is a third of Latvia's population. The city lies on the Gulf of Riga at the mouth of the Daugava river where it meets the Ba ...
to aid the city after Napoleon's armies had laid siege to it. As he attempted to help the sick and wounded, he realized that his education at Dorpat had been inadequate, and upon his graduation, he notified his father that he would need to go abroad to "finish" his education. In his autobiography, his discontent with his education at Dorpat inspired him to write a lengthy appraisal of education in general, a summary that dominated the content of the book. After leaving Tartu, he continued his education in
Berlin Berlin ( , ) is the capital and largest city of Germany by both area and population. Its 3.7 million inhabitants make it the European Union's most populous city, according to population within city limits. One of Germany's sixteen constitue ...
,
Vienna en, Viennese , iso_code = AT-9 , registration_plate = W , postal_code_type = Postal code , postal_code = , timezone = CET , utc_offset = +1 , timezone_DST ...
, and
Würzburg Würzburg (; Main-Franconian: ) is a city in the region of Franconia in the north of the German state of Bavaria. Würzburg is the administrative seat of the ''Regierungsbezirk'' Lower Franconia. It spans the banks of the Main River. Würzburg is ...
, where
Ignaz Döllinger Ignaz Döllinger (27 May 1770 – 14 January 1841) was a German doctor, anatomist and physiologist and one of the first professors to understand and treat medicine as a natural science. Biography Ignaz Döllinger was born in 1770 in Bamberg, w ...
introduced him to the new field of
embryology Embryology (from Greek ἔμβρυον, ''embryon'', "the unborn, embryo"; and -λογία, '' -logia'') is the branch of animal biology that studies the prenatal development of gametes (sex cells), fertilization, and development of embryos and ...
. In 1817, he became a professor at
Königsberg University Königsberg (, ) was the historic Prussian city that is now Kaliningrad, Russia. Königsberg was founded in 1255 on the site of the ancient Old Prussian settlement ''Twangste'' by the Teutonic Knights during the Northern Crusades, and was named ...
and full professor of
zoology Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...
in 1821, and of
anatomy Anatomy () is the branch of biology concerned with the study of the structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having its ...
in 1826. In 1829, he taught briefly in
St Petersburg Saint Petersburg ( rus, links=no, Санкт-Петербург, a=Ru-Sankt Peterburg Leningrad Petrograd Piter.ogg, r=Sankt-Peterburg, p=ˈsankt pʲɪtʲɪrˈburk), formerly known as Petrograd (1914–1924) and later Leningrad (1924–1991), i ...
, but returned to
Königsberg Königsberg (, ) was the historic Prussian city that is now Kaliningrad, Russia. Königsberg was founded in 1255 on the site of the ancient Old Prussian settlement ''Twangste'' by the Teutonic Knights during the Northern Crusades, and was named ...
(Kaliningrad). In 1834, Baer moved back to St Petersburg and joined the
St Petersburg Academy of Sciences The Russian Academy of Sciences (RAS; russian: Росси́йская акаде́мия нау́к (РАН) ''Rossíyskaya akadémiya naúk'') consists of the national academy of Russia; a network of scientific research institutes from across t ...
, first in zoology (1834–46) and then in
comparative anatomy Comparative anatomy is the study of similarities and differences in the anatomy of different species. It is closely related to evolutionary biology and phylogeny (the evolution of species). The science began in the classical era, continuing in t ...
and
physiology Physiology (; ) is the scientific study of functions and mechanisms in a living system. As a sub-discipline of biology, physiology focuses on how organisms, organ systems, individual organs, cells, and biomolecules carry out the chemical ...
(1846–62). His interests while there were anatomy,
ichthyology Ichthyology is the branch of zoology devoted to the study of fish, including bony fish ( Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of Octob ...
,
ethnography Ethnography (from Greek ''ethnos'' "folk, people, nation" and ''grapho'' "I write") is a branch of anthropology and the systematic study of individual cultures. Ethnography explores cultural phenomena from the point of view of the subject o ...
,
anthropology Anthropology is the scientific study of humanity, concerned with human behavior, human biology, cultures, societies, and linguistics, in both the present and past, including past human species. Social anthropology studies patterns of behavi ...
, and
geography Geography (from Greek: , ''geographia''. Combination of Greek words ‘Geo’ (The Earth) and ‘Graphien’ (to describe), literally "earth description") is a field of science devoted to the study of the lands, features, inhabitants, and ...
. While embryology had kept his attention in Königsberg, then in Russia von Baer engaged in a great deal of field research, including the exploration of the island
Novaya Zemlya Novaya Zemlya (, also , ; rus, Но́вая Земля́, p=ˈnovəjə zʲɪmˈlʲa, ) is an archipelago in northern Russia. It is situated in the Arctic Ocean, in the extreme northeast of Europe, with Cape Flissingsky, on the northern island, ...
. The last years of his life (1867–76) were spent in Dorpat, where he became a major critic of
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended fr ...
.


Contributions


Embryology

von Baer studied the embryonic development of animals, discovering the
blastula Blastulation is the stage in early animal embryonic development that produces the blastula. In mammalian development the blastula develops into the blastocyst with a differentiated inner cell mass and an outer trophectoderm. The blastula (from ...
stage of development and the
notochord In anatomy, the notochord is a flexible rod which is similar in structure to the stiffer cartilage. If a species has a notochord at any stage of its life cycle (along with 4 other features), it is, by definition, a chordate. The notochord consis ...
. Together with
Heinz Christian Pander Heinz Christian Pander, also Christian Heinrich Pander ( – ), was a Russian Empire ethnic Baltic German biologist and embryologist. Biography In 1817 he received his doctorate from the University of Würzburg, and spent several years (1827 ...
and based on the work by
Caspar Friedrich Wolff Caspar Friedrich Wolff (18 January 1733 – 22 February 1794) was a German physiologist and one of the founders of embryology. Life Wolff was born in Berlin, Brandenburg. In 1759 he graduated as an M.D. from the University of Halle with his disse ...
, he described the
germ layer A germ layer is a primary layer of cells that forms during embryonic development. The three germ layers in vertebrates are particularly pronounced; however, all eumetazoans (animals that are sister taxa to the sponges) produce two or three pr ...
theory of development (
ectoderm The ectoderm is one of the three primary germ layers formed in early embryonic development. It is the outermost layer, and is superficial to the mesoderm (the middle layer) and endoderm (the innermost layer). It emerges and originates from t ...
,
mesoderm The mesoderm is the middle layer of the three germ layers that develops during gastrulation in the very early development of the embryo of most animals. The outer layer is the ectoderm, and the inner layer is the endoderm.Langman's Medical E ...
, and
endoderm Endoderm is the innermost of the three primary germ layers in the very early embryo. The other two layers are the ectoderm (outside layer) and mesoderm (middle layer). Cells migrating inward along the archenteron form the inner layer of the gast ...
) as a principle in a variety of species, laying the foundation for comparative embryology in the book ''Über Entwickelungsgeschichte der Thiere'' (1828). In 1826, Baer discovered the mammalian
ovum The egg cell, or ovum (plural ova), is the female reproductive cell, or gamete, in most anisogamous organisms (organisms that reproduce sexually with a larger, female gamete and a smaller, male one). The term is used when the female gamete is ...
. The human ovum was first described by
Edgar Allen Edgar Allen (May 2, 1892 – February 3, 1943) was an American anatomist and physiologist. He is known for the discovery of estrogen and his role in creating the field of endocrinology. Born on Cañon (Canyon) City, Colorado, Allen was educate ...
in 1928. In 1827, he completed research ''Ovi Mammalium et Hominis genesi'' for St Petersburg's Academy of Science (published at Leipzig). In 1827 von Baer became the first person to observe human
ova , abbreviated as OVA and sometimes as OAV (original animation video), are Japanese animated films and series made specially for release in home video formats without prior showings on television or in theaters, though the first part of an OVA s ...
. Only in 1876 did
Oscar Hertwig Oscar Hertwig (21 April 1849 in Friedberg – 25 October 1922 in Berlin) was a German embryologist and zoologist known for his research in developmental biology and evolution. Hertwig is credited as the first man to observe sexual reproduction ...
prove that fertilization is due to fusion of an egg and sperm cell. von Baer formulated what became known as Baer's laws of embryology: # General characteristics of the group to which an embryo belongs develop before special characteristics. # General structural relations are likewise formed before the most specific appear. # The form of any given embryo does not converge upon other definite forms, but separates itself from them. # The embryo of a higher animal form never resembles the adult of another animal form, such as one less evolved, but only its embryo.


Permafrost research

Baer was a genius scientist covering not only the topics of embryology and ethnology, he also was especially interested in the geography of the northern parts of Russia, and explored
Novaya Zemlya Novaya Zemlya (, also , ; rus, Но́вая Земля́, p=ˈnovəjə zʲɪmˈlʲa, ) is an archipelago in northern Russia. It is situated in the Arctic Ocean, in the extreme northeast of Europe, with Cape Flissingsky, on the northern island, ...
in 1837. In these
arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
environments, he was studying
periglacial Periglaciation (adjective: "periglacial", also referring to places at the edges of glacial areas) describes geomorphic processes that result from seasonal thawing of snow in areas of permafrost, the runoff from which refreezes in ice wedges and ot ...
features,
permafrost Permafrost is ground that continuously remains below 0 °C (32 °F) for two or more years, located on land or under the ocean. Most common in the Northern Hemisphere, around 15% of the Northern Hemisphere or 11% of the global surface ...
occurrences, and collecting biological specimens. Other travels led him to
subarctic The subarctic zone is a region in the Northern Hemisphere immediately south of the true Arctic, north of humid continental regions and covering much of Alaska, Canada, Iceland, the north of Scandinavia, Siberia, and the Cairngorms. Generally, ...
regions of the North Cape and Lapland, but also to the
Caspian Sea The Caspian Sea is the world's largest inland body of water, often described as the world's largest lake or a full-fledged sea. An endorheic basin, it lies between Europe and Asia; east of the Caucasus, west of the broad steppe of Central Asia ...
. He was one of the founders of the
Russian Geographical Society The Russian Geographical Society (russian: Ру́сское географи́ческое о́бщество «РГО»), or RGO, is a learned society based in Saint Petersburg, Russia. It promotes geography, exploration and nature protection wi ...
. Thanks to Baer's research expeditions, the scientific investigation of permafrost began in Russia. Baer recorded the importance of permafrost research even before 1837 when observing in detail the
geothermal gradient Geothermal gradient is the rate of temperature change with respect to increasing depth in Earth's interior. As a general rule, the crust temperature rises with depth due to the heat flow from the much hotter mantle; away from tectonic plate b ...
from a 116.7 m deep shaft in
Yakutsk Yakutsk (russian: Якутск, p=jɪˈkutsk; sah, Дьокуускай, translit=Djokuuskay, ) is the capital city of the Sakha Republic, Russia, located about south of the Arctic Circle. Fueled by the mining industry, Yakutsk has become one of ...
. At the end of the 1830s, he recommended sending expeditions to explore permafrost in Siberia and suggested
Alexander von Middendorff Alexander Theodor von Middendorff (russian: Алекса́ндр Фёдорович Ми́ддендорф; tr. ; 18 August 1815 – 24 January 1894) was a zoologist and explorer of Baltic German and Estonian extraction. He is known for his ex ...
as leader. Baer's expedition instructions written for Middendorff comprised over 200 pages. Baer summarized his knowledge in 1842/43 in a print-ready typescript. The German title is „Materialien zur Kenntniss des unvergänglichen Boden-Eises in Sibirien“ (=materials for the knowledge of the perennial ground ice in Siberia). This world's first permafrost textbook was conceived as a complete work for printing. But it remained lost for more than 150 years. However, from 1838 onwards, Baer published a larger number of small publications on permafrost. Numerous of Baer's papers on permafrost were already published as early as 1837 and 1838. Well known was his paper "On the Ground Ice or Frozen Soil of Siberia", published in the Journal of the Royal Geographical Society of London (1838, pp. 210–213) and reprinted 1839 in the American Journal of Sciences and Arts by S. Silliman. There are many other publications and small notes on permafrost by Baer, as shown in the Karl Ernst von Baer museum in
Tartu Tartu is the second largest city in Estonia after the Northern European country's political and financial capital, Tallinn. Tartu has a population of 91,407 (as of 2021). It is southeast of Tallinn and 245 kilometres (152 miles) northeast of ...
(
Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, a ...
), now part of the
Estonian University of Life Sciences The Estonian University of Life Sciences ( Estonian: ''Eesti Maaülikool'', EMÜ) located in Tartu, Estonia, is the former Estonian Agricultural University, which was established in 1951 and renamed and restructured in November 2005. Eesti Maa ...
. There are quite a number of studies in Russian about the origin of permafrost research. Russian authors usually relate with it the name
Alexander von Middendorff Alexander Theodor von Middendorff (russian: Алекса́ндр Фёдорович Ми́ддендорф; tr. ; 18 August 1815 – 24 January 1894) was a zoologist and explorer of Baltic German and Estonian extraction. He is known for his ex ...
(1815–1894), as he did much scientific work during the years 1842–1845 concerning permafrost on
Taimyr Peninsula The Taymyr Peninsula (russian: Таймырский полуостров, Taymyrsky poluostrov) is a peninsula in the Far North of Russia, in the Siberian Federal District, that forms the northernmost part of the mainland of Eurasia. Administrati ...
and in East-
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
. However, Russian scientists during the 1940s also realized, that it was K. E. Baer who initiated this expedition and that the origin of scientific permafrost research must be fixed with Baer's thorough earlier scientific work. They even believed, that the scepticism about the permafrost findings and publications of Middendorff would not have risen, if Baer's original "materials for the study of the perennial ground-ice" would have been published in 1842 as intended. This was realized also by the Russian Academy of Sciences that honoured Baer with the publication of a tentative Russian translation done already in 1842 by Sumgin. These facts were completely forgotten until after the Second World War. In North America, permafrost research started after the Second World War with the creation of the
Cold Regions Research and Engineering Laboratory The Cold Regions Research and Engineering Laboratory (CRREL) is a United States Army Corps of Engineers, Engineer Research and Development Center research facility headquartered in Hanover, New Hampshire, that provides scientific and engineering ...
(CRREL), a division of the US army. It was realized that the understanding of frozen ground and permafrost are essential factors in strategic northern areas during the
Cold War The Cold War is a term commonly used to refer to a period of geopolitical tension between the United States and the Soviet Union and their respective allies, the Western Bloc and the Eastern Bloc. The term '' cold war'' is used because the ...
. In the Soviet Union, the
Melnikov Permafrost Institute Melnikov Permafrost Institute of the Siberian Branch of the Russian Academy of Science (russian: Институт мерзлотоведения имени П. И. Мельникова СО РАН) is a research institute based in Yakutsk, Russia, ...
in
Yakutsk Yakutsk (russian: Якутск, p=jɪˈkutsk; sah, Дьокуускай, translit=Djokuuskay, ) is the capital city of the Sakha Republic, Russia, located about south of the Arctic Circle. Fueled by the mining industry, Yakutsk has become one of ...
had similar aims. The first post-World War major contact between groups of senior Russian and American frozen ground researchers took place in November 1963 in Yakutsk.However, Baer's permafrost textbook remained still undiscovered. Thus in 2001 the discovery and annotated publication of the typescript from 1843 in the library
archive An archive is an accumulation of historical records or materials – in any medium – or the physical facility in which they are located. Archives contain primary source documents that have accumulated over the course of an individual or ...
s of the
University of Giessen University of Giessen, official name Justus Liebig University Giessen (german: Justus-Liebig-Universität Gießen), is a large public research university in Giessen, Hesse, Germany. It is named after its most famous faculty member, Justus von L ...
was a scientific sensation. The full text of Baer's work is available online (234 pages). The editor Lorenz King added to the
facsimile A facsimile (from Latin ''fac simile'', "to make alike") is a copy or reproduction of an old book, manuscript, map, Old master print, art print, or other item of historical value that is as true to the original source as possible. It differs from ...
reprint a preface in English, two colour permafrost maps of Eurasia and some figures of permafrost features. Baer's text is introduced with detailed comments and references on additional 66 pages written by the
Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, a ...
n historian Erki Tammiksaar. The work is fascinating to read, because both Baer's observations on permafrost distribution and his periglacial morphological descriptions are largely still correct today. He distinguished between "continental" and "insular" permafrost, saw the temporary existence of permafrost and postulated the formation and further development of permafrost as a result of the complex physio-geographical,
geological Geology () is a branch of natural science concerned with Earth and other astronomical objects, the features or rocks of which it is composed, and the processes by which they change over time. Modern geology significantly overlaps all other Eart ...
and
floristic {{Short pages monitor


External links


Bibliography and links
at
Max Planck Institute Max or MAX may refer to: Animals * Max (dog) (1983–2013), at one time purported to be the world's oldest living dog * Max (English Springer Spaniel), the first pet dog to win the PDSA Order of Merit (animal equivalent of OBE) * Max (gorilla) (1 ...

Overview of Piibe (Piep) manor (with photo of memorial stone)



Glossary of Permafrost and Related Ground-Ice Terms (IPA)

International Conferences on Permafrost



Ajaloolane: Karl Ernst von Baer oli Venemaa luure asutaja
(in Estonian) {{DEFAULTSORT:Baer, Karl Ernst Von 1792 births 1876 deaths People from Väike-Maarja Parish People from Kreis Jerwen Baltic-German people Estonian explorers German explorers Explorers from the Russian Empire Biologists from the Russian Empire German embryologists Orthogenesis Proto-evolutionary biologists Recipients of the Copley Medal Novaya Zemlya Explorers of the Arctic 19th-century Estonian people Estonian nobility German knights University of Tartu alumni University of Königsberg faculty Members of the Royal Netherlands Academy of Arts and Sciences Members of the Royal Swedish Academy of Sciences Fellows of the American Academy of Arts and Sciences Founding members of the Russian Geographical Society Full members of the Saint Petersburg Academy of Sciences Honorary members of the Saint Petersburg Academy of Sciences Foreign Members of the Royal Society Foreign associates of the National Academy of Sciences German military personnel of the Napoleonic Wars Recipients of the Pour le Mérite (civil class) Burials at Raadi cemetery Members of the Göttingen Academy of Sciences and Humanities